Rose and Myrtle Sylvester look up to their older sister, Peggy. She is the sensible, reliable one in the household of women headed by their grandmother, Grace Booth, and their mother, Mary Sylvester. When war is declared in 1939 they must face the hardships together and huge changes in their lives are inevitable. For Rose, there is the chance to fulfil her dream of becoming a clippie on Sheffield's trams like Peggy. But for Myrtle, the studious, clever one in the family, war may shatter her ambitions.

When the tram on which Peggy is a conductress is caught in a bomb blast, she bravely helps to rescue her passengers. One of them is a young soldier, Terry Price, and he and Peggy begin courting. They meet every time he can get leave, but eventually Terry is posted abroad and she hears nothing from him. Worse still, Peggy must break the devastating news to her family that she is pregnant.

The shock waves that ripple through the family will affect each and every one of them and life will never be the same again.